Drawing on historical perspectives, personal excursions, and decades of professional research and work in the field, Paul Schullery illuminates many of the possible truths embedded within the natural and cultural reality that is Yellowstone National Park. By varying the scale of observation—from a single encounter between a cow elk and a grizzly bear to the sweeping forces of evolution—Schullery celebrates the park’s history and future potential as a laboratory of ideas. It is, as he states, a place with “layers of meaning waiting to be explored . . . many possible truths to be weighed.” He thus invites us all to participate in the “Yellowstone conversation.” According to Schullery, national parks allow for the study of relatively unmanipulated ecological processes even amidst civilization’s increasing influence. They act as reservoirs for water, wildlife, and essential wildness. The uncertainties inherent in wild landscapes and in the unfolding idea of Yellowstone allow scholarly and popular dialogues to advance management practices and public understanding. Through this inquiry, Schullery establishes a framework for approaching the conservation and the experience of America’s great wildlands. Paul Schullery delivered this lecture on March 26, 2014, at the 19th annual symposium sponsored by the Wallace Stegner Center for Land, Resources and the Environment at the S. J. Quinney College of Law of the University of Utah.